Chrome and Firefox can now talk to each other via WebRTC. It's a new set of technologies that brings clear crisp voice, sharp high-definition (HD) video and low-delay communication to the web browser without any plug-in.
To try this yourself, you’ll need desktop Chrome 25 Beta and Firefox Nightly for Desktop. In Firefox, you'll need to go to about:config and set the media.peerconnection.enabled pref to "true”. Then head over to the WebRTC demo site and start calling.
Read more on Chrome and Firefox blogs:
Hello Firefox, this is Chrome calling!
Hello Chrome, it’s Firefox calling!
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A conversation Between Chrome and Firefox - WebRTC
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